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Complete Publications


















2024

  • Schools M, Hedges SB. 2024. A new forest lizard fauna from Caribbean islands (Squamata, Diploglossidae, Celestinae). Zootaxa 5554: 1-306. (Request from author)
  • Craig JM, Hedges SB, Kumar S. 2024. Completing a molecular timetree of primates. Frontiers in Bioinformatics 4:1495417. (Open Access)
  • Jesse WAM, Ellers J, Behm JE, Costa GC, Hedges SB, Helmus MR. 2024. Elevated human impact on islands increases the introduction and extinction status of native insular reptiles. Ecography e06817. (Open Access)
  • Ali J, Hedges SB. 2024. Land bridges and rafting theories to explain terrestrial-vertebrate biodiversity on Madagascar. Annual Review of Marine Science Vol. 17 (Early Publication): DOI: 10.1146/annurev-marine-032223-025654 (Request from author)
  • Ali J, Hedges SB. 2024. Paleogeography of the Aves Ridge and its potential role as a bio-colonization pathway linking South America and the Greater Antilles in the mid-Cenozoic. Earth-Science Reviews: DOI: doi.org/10.1016/j.earscirev.2024.104823 (Request from author)

2023

  • Craig JM, Bamba GL, Barba J, Hedges SB, Kumar S. 2023. Completing a molecular timetree of apes and monkeys. Frontiers in Bioinformatics DOI: 10.3389/fbinf.2023.1284744. (Open Access)
  • Luedtke J, et al. 2023. Ongoing declines for the world's amphibians in the face of emerging threats. Nature 622:308–314. (Request from author)
  • Craig J, Kumar S, Hedges SB. 2023. The origin of eukaryotes and rise in complexity were synchronous with the rise in oxygen. Frontiers in Bioinformatics DOI: 10.3389/fbinf.2023.1233281. (Request from author)
  • Hedges SB. 2023. The conservation and biogeography of amphibians in the Caribbean. Phyllomedusa 22: 87-93. (Request from author)
  • Means DB, Heinicke MP, Hedges SB, MacCulloch RD, Lathrop A. 2023. Exceptional diversity of Pristimantis landfrogs (Anura: Terraranae) on the Wokomung Massif, Guyana, with descriptions of three new species. Journal of Vertebrate Biology 72: 23026.1-26 (Open Access)
  • Ali J, Hedges SB. 2023. The colonisation of Madagascar by land-bound vertebrates. Biological Reviews 98(5): 1583-1606. (Request from author)
  • Stanojevic M, Andjelkovic J, Kasprowicz A, Huuki LA, Chao J, Hedges SB, Kumar S, Obradovic Z. 2023. Discovering research articles containing evolutionary timetrees by machine learning. Bioinformatics (Article Link)

2022

  • Hedges SB. 2022. A new species of frog from the Caribbean island of Montserrat (Eleutherodactylidae, Eleutherodactylus). Zootaxa 5219: 375-387. (Request from author)
  • Landestoy MA, Schools M, Hedges SB. 2022. A new genus and species of Caribbean forest lizard (Diploglossidae; Celestinae)from southern Hispaniola. Zootaxa 5219: 201-226. (Request from author)
  • Kumar S, Suleski M, Craig JM, Kasprowicz AE, Sanderford M, Li M, Stecher G., Hedges SB. 2022. TimeTree 5: an expanded resource for species divergence times. Mol. Biol. Evol.. (Open Access Link)
  • Schools M, Kasprowicz A, Hedges SB. 2022. Phylogenomic data resolve the historical biogeography and ecomorphs of Neotropical forest lizards (Squamata, Diploglossidae). Molecular Phylogenetics and Evolution. (Request from author)
  • Ali J, Hedges SB. 2022. A review of geological evidence bearing on proposed Cenozoic land connections between Madagascar and Africa and its relevance to biogeography. Earth-Science Reviews. Published online 7 July 2022 (Request from author)
  • Yuan ML, Frederick JH, McGuire JA, Bell RC, Smith SR, Fenton C, Cassius J, Williams R, Wang IJ, Powell R, Hedges SB. 2022 Endemism, invasion, and overseas dispersal: the phylogeographic history of the Lesser Antillean frog, Eleutherodactylus johnstonei. Biological Invasions 24:2702-2722. Published online 7 May 2022 (Open Access Link)
  • Cox N, Young BE, Bowles P, Fernandez M, Marin J, Rapacciuolo G, Bohm M, Brooks TM, Hedges SB, et al. 2022. A global reptile assessment highlights shared conservation needs of tetrapods. Nature. (Open Access Link)
  • Craig JM, Kumar S, Hedges SB. 2022. Limitations of phylogenomic data can drive inferred speciation rate shifts. Mol. Biol. Evol. 39: msac038. (Open Access Link)

2021

  • Diaz LM, Fong A, Salas D, Hedges SB. 2021. A new semifossorial snake of the genus Arrhyton (Squamata: Dipsadidae) from eastern Cuba, with taxonomic comments on other species. Zootaxa 5052: 406-418. (Request from author)
  • Ali JR, Hedges SB. 2021. Colonizing the Caribbean: New geological data and an updated land-vertebrate colonization record challenge the GAARlandia land-bridge hypothesis. J. Biogeography https://doi.org/10.1111/jbi.14234. E-print
  • Moore RD, Hedges SB. 2021. Private conservation of remnant forest ecosystems to support sustainable development: Grand Bois privately protected area, Haiti. Pp. 50-53 in Kettunen M, Dudley N, Gorricho J, Hickey V, Krueger L, MacKinnon K, Oglethorpe J, Paxton M, Robinson JG, Sekhran N, Building on nature: area-based conservation as a key tool for delivering SDGs (IEEP, IUCN WCPA, The Nature Conservancy, The World Bank, UNDP, Wildlife Conservation Society and WWF). (Request from author)
  • Schools M, Hedges SB. 2021. Phylogenetics, classification, and biogeography of the Neotropical forest lizards (Squamata, Diploglossidae). Zootaxa 4974:201-257. (Request from author)
  • Su D, Yang L, Shi X, Ma X, Zhou X, Hedges SB, Zhong B. 2021. Large-scale phylogenomic analyses reveal the monophyly of bryophytes and Neoproterozoic origin of land plants. Mol. Biol. Evol. https://doi.org/10.1093/molbev/msab106. (published online). E-print
  • Chapple DG, Roll U, Bohm M, Aguilar R, Amey AP, Austin CC, Baling M, Barley AJ...Hedges, SB...et al. 2021. Conservation status of the world’s skinks (Scincidae): taxonomic and geographic patterns in extinction risk. Conservation Biology (published online). (Request from author)

2020

  • Köhler G, Hedges SB. 2020. A replacement name for the Hispaniolan anole formerly referred to as Anolis chlorocyanus Duméril & Bibron, 1837. Caribbean Herpetology, 70:1-3. E-print
  • McCranie JR, Matthews AJ, Hedges SB. 2020. A morphological and molecular revision of lizards of the genus Marisora (Squamata: Mabuyidae) from Central America and Mexico, with descriptions of four new species. Zootaxa 4763: 301-353. (Request from author)
  • Hedges SB. 2020. Molecular clocks, flotsam, and Caribbean islands. Pp. 349–351 in Cox, CB, Laddle RJ, & Moore PD, Biogeography: an ecological and evolutionary Approach. 10th Edition (John Wiley & Sons, Inc., Hoboken, New Jersey). (Request from author)

2019

  • Köhler G, Zimmer C, McGrath K, Hedges SB. 2019. A revision of the genus Audantia of Hispaniola with description of four new species (Reptilia, Squamata, Dactyloidae). Novitates Caribaea. 14: 1-104. doi/10.33800/nc.v0i14.201 E-print (6MB) E-print (55MB)
  • Hedges SB, Powell R, Henderson RW, Hanson S, and Murphy JC. 2019. Definition of the Caribbean Islands biogeographic region, with checklist and recommendations for standardized common names of amphibians and reptiles. Caribb. Herpetol. 67:1-53. doi/10.31611/ch.67 E-print
  • Hedges SB, Cohen WB, Timyan J, Yang Z. 2018. Reply to Wampler et al.: Deforestation and biodiversity loss should not be sugarcoated. Proc. Natl. Acad Sci. USA. 116: doi/10.1073/pnas.1901879116. (Request from author)
  • Hedges SB, Lorvelec O, Barré N, Vidal N, Pavis C. 2019. On the taxonomic recognition of skinks from the Guadeloupe Archipelago (Squamata, Mabuyidae, Mabuya). Caribbean Herpetology, 64:1-7. E-print

2018

  • Rapacciuolo G, Graham CH, Marin J, Behm JE, Costa GC, Hedges SB, Helmus MR, Radeloff VC, Young BE, Brooks TM. 2018. Species diversity as a surrogate for conservation of phylogenetic and functional diversity in terrestrial vertebrates across the Americas. Nature Ecol. Evol. 2(12):10.1038/s41559-018-0744-7 (Request from author)
  • Hedges SB, Cohen WB, Timyan J, Yang Z. 2018. Haiti's biodiversity threatened by nearly complete loss of primary forest. Proc. Natl. Acad Sci. USA. 115(46):11850-11855. (Request from author)
  • Schmid M, Bogart JP, Hedges SB. (eds) 2018. The arboranan frogs: evolution, biology, and cytogenetics. Cytogenet Genome Res 155, No. 1–4. 326 p., 956 fig.
  • Hedges SB, Tao Q, Walker M, Kumar S. 2018. Accurate timetrees require accurate calibrations. Proc. Natl. Acad Sci. USA. 115(41):E9510-E9511. (Request from author)
  • Miralles A, Marin J, Markus D, Herrel A, Hedges SB, Vidal N. 2018. Molecular evidence for the paraphyly of Scolecophidia and its evolutionary implications. J Evol Biol. doi: 10.1111/jeb.13373. (Request from author)
  • Marin J, Hedges SB. 2018. Undersampling genomes has biased time and rate estimates throughout the tree of life. Mol. Biol. Evol. 35: 2077–2084. (Request from author)
  • Landestoy MA, Turner DB, Marion AB, Hedges SB. 2018. A new species of Caribbean toad (Bufonidae, Peltophryne) from southern Hispaniola. Zootaxa, published online 5 April 2018: 10.11646/zootaxa.4403.3.6. (Request from author)
  • Marin J, Rapacciuolo G, Costa GC, Graham CH, Brooks TM, Young BE, Radeloff VC, Behm JE, Helmus MR, Hedges SB. 2018. Evolutionary time drives global tetrapod diversity. Roy Soc. Proc B, published online 7 February 2018. doi: 10.1098/rspb.2017.2378. (Request from author)

2017

  • Holt BG, Costa GC, Penone C, Lessard J-P, Brooks TM, Davidson AD, Hedges SB, Radeloff VC, Rahbek C, Rondinini C, Graham CH. 2017. Environmental variation is a major predictor of global trait turnover in mammals. J. Biogeography 2017: 1-13. (Request from author)
  • Heinicke MP, Lemmon AR, Lemmon EM, McGrath K, Hedges SB. 2017. Phylogenomic support for evolutionary relationships of New World direct-developing frogs (Anura: Terraranae). Molec. Phylogenet. Evol. 118: 145-155. (Request from author)
  • Lorvelec O, Barré N, Chalifour J, Teynié A, Pisanu B, Hedges SB. 2017. Discovery of a population of Spondylurus powelli (Squamata: Mabuyidae) on Île Tintamarre (Saint-Martin, French Antilles) and comments on relationships among skinks of the Anguilla Bank. Caribbean Herpetology, 59: 1–8. E-print
  • Brum FT, Graham CH, Costa GC, Hedges SB, Penone C, Radeloff VC, Rondinini C, Loyola R, Davidson AD. 2017. Global priorities for conservation across multiple dimensions of mammalian diversity. Proc. Natl. Acad. Sci. USA. 114(29): 7641–7646. (Request from author)
  • Kumar S, Stecher G, Suleski M, Hedges SB. 2017. TimeTree: A resource for timelines, timetrees, and divergence times. Mol. Biol. Evol. doi: 10.1093/molbev/msx116. (Request from author)
  • Tucker DB, Hedges SB, Colli GR, Pyron RA, Sites JW, Jr. 2017. Genomic timetree and historical biogeography of Caribbean island ameiva lizards (Pholidoscelis: Teiidae). Ecology and Evolution. doi: 10.1002/ece3.3157. (Request from author)
  • Rapacciuolo G, Marin J, Costa GC, Helmus MR, Behm JE, Brooks TM, Hedges SB, Radeloff VC, Young BE, Graham CH. 2017. The signature of human pressure history on the biogeography of body mass in tetrapods. Global Ecology and Biogeography. doi: 10.1111/geb.12612. (Request from author)
  • Marin J, Battistuzzi FU, Brown AC, Hedges SB. 2017. The Timetree of prokaryotes: new insights into their evolution and speciation. Mol. Biol. Evol. doi: 10.1093/molbev/msw245. (Request from author)

2016

  • Penone C, Weinstein BG, Graham CH, Brooks TM, Rondinini C, Hedges SB, Davidson AD, Costa GC. 2016. Global mammal beta diversity shows parallel assemblage structure in similar but isolated environments. Proc. R. Soc. B 283: 20161028. doi: 10.1098/rspb.2016.1028. (Request from author)
  • Tucker DB, Colli GR, Giugliano LG, Hedges SB, Hendry CR, Lemmon EM, Lemmon AR, Sites JW Jr, Pyron RA. Methodological congruence in phylogenomic analyses with morphological support for teiid lizards (Sauria: Teiidae). Molec. Phylogenet. Evol. 103: 75-84. E-print
  • Mahler DL, Lambert SM, Geneva AJ, Ng J, Hedges SB, Losos JB, Glor RE. 2016. Discovery of a giant chameleon-like lizard (Anolis) on Hispaniola and its significance to understanding replicated adaptive radiations. The American Naturalist 188(3): doi: 10.1086/687566. E-print
  • Köhler G, Rodríguez MJ, Hedges SB. 2016. A new dune-dwelling lizard of the genus Leiocephalus (Iguania, Leiocephalidae) from the Dominican Republic. Zootaxa 4121: 517-532. E-print
  • Duellman WE, Marion AB, Hedges SB. 2016. Phylogenetics, classification, and biogeography of the treefrogs (Amphibia: Anura: Arboranae). Zootaxa 4104: 1-109. (Request from author)
  • Marin J, Hedges SB. 2016. Time best explains global variation in species richness of amphibians, birds and mammals. J. Biogeography 43: 1069-1079. (Request from author)
  • Hedges SB, Lorvelec O, Barré N, Berchel J, Combot MD, Vidal N, Pavis C. 2016. A new species of skink from the Guadeloupe Archipelago (Squamata, Mabuyidae, Mabuya). Caribbean Herpetology 53:1-14. E-print
  • Kumar S, Hedges SB. 2016. Advances in time estimation methods for molecular data. Mol. Biol. Evol. 33: 863-869. E-print
  • Köhler G, Hedges SB. 2016. A revision of the green anoles of Hispaniola with description of eight new species (Reptilia, Squamata, Dactyloidae). Novitates Caribaea 9: 1-135. E-print (Low-res, 5MB) E-print (Hi-res, 70MB) KEY
  • Angulo A, May RV, Icochea J, Chaparro JC, Hedges SB, Lehr E, López DG, Lundberg M, & Moravec J. 2016. Parque Nacional Yanachaga-Chemillén, Pasco, Perú. Anfibios del bosque de nubes y su zona de amortiguamiento. Field Guides 738: 1-6. E-print
  • McCranie JR, Hedges SB. 2016. Molecular phylogeny and taxonomy of the Epictia goudotii Species complex (Serpentes: Leptotyphlopidae: Epictinae) in Middle America and northern South America. PeerJ 4:e1551; doi 10.7717/peerj.1551. E-print

2015

  • Nagy ZT, Marion AB, Glaw F, Miralles A., Noppers J, Vences M, Hedges SB. 2015. Molecular systematics and undescribed diversity of Madagascan scolecophidian snakes (Squamata: Serpentes). Zootaxa 4040: 031–047. (Request from author)
  • Schmid M et al. (104 authors) 2015. Third report on chicken genes and chromosomes. Cytogenet Genome Res. 145:78-179 (Request from author)
  • Heinicke MP, Barrio-Amorós CL, and SB Hedges. 16 Jul 2015. Molecular and morphological data support recognition of a new genus of New World direct-developing frog (Anura: Terrarana) from an under-sampled region of South America​. Zootaxa 3986 (2): 151–172. (Request from author)
  • Dίaz, LM, and SB Hedges. 2015. Another new cryptic frog related to Eleutherodactylus varleyi Dunn (Amphibia: Anura: Eleutherodactylidae), from eastern Cuba​. Solenodon 12:124-135. E-print
  • Köhler G, Hedges SB. 2015. Anolis chlorocyanus Duméril & Bibron, 1837 and Anolis coelestinus Cope, 1862 (Reptilia, Squamata): proposed conservation of the specific names and designation of a neotype for A. chlorocyanus. Bull. Zool. Nomenclature 72: 45-49. E-print
  • Hedges, S. B., J. Marin, M. Suleski, M. Paymer & S. Kumar. 2015. Tree of Life Reveals Clock-Like Speciation and Diversification. Mol. Biol. Evol. 32(4):835–845 E-print, Supplement 1, Supplement 2

2014

  • Schmid M, Steinlein C, Feichtinger W, Haaf T, Mijares-Urrutia A, Schargel WE, Hedges SB. 21 Oct 2014. Cytogenetic studies on Gonatodes (Reptilia, Squamata, Sphaerodactylidae).Cytogenet Genome Res. 144(1):47-61. E-Print
  • Hedges SB. 2014. The high-level classification of skinks (Reptilia, Squamata, Scincomorpha). Zootaxa 3765: 317-338. (Request from author)
  • Hedges SB, Marion AB, Lipp KM, Marin J, Vidal N. 2014. A taxonomic framework for typhlopid snakes from the Caribbean and other regions (Reptilia, Squamata). Caribbean Herpetology 49:1-61. E-print
  • Hedges SB. 2014. The origin of life. In Oxford Bibliographies in Evolutionary Biology Ed. Jonathan Losos. New York: Oxford University Press. Oxford Bibliographies website. (Request from author)

2013

  • Wegener, JE, Swoboda S, Hawlitschek O, Franzen M, Wallach V, Vences M, Nagy Z, Hedges SB, Kohler J, Glaw F. 2013. Morphological variation and taxonomic assessment of the endemic Malagasy blind snake family Xenotyphlopidae. Spixiana 36: 269-282. E-print
  • McCranie JR, Hedges SB. 2013. A review of the Cnemidophorus lemniscatus group in Central America (Squamata: Teiidae), with comments on other species in the group.Zootaxa 3722: 301-316. Request from author
  • Marin J, Donnellan SC, Hedges SB, Puillandre N, Aplin KP, Doughty P, Hutchinson MN, Couloux A, Vidal N. 2013. Hidden species diversity of Australian burrowing snakes (Ramphotyphlops). Biol. J. Linn. Soc. 110: 427-441. E-print
  • McCranie JR, Hedges SB. 2013. A new species of Phyllodactylus (Reptilia, Squamata, Gekkonoidea, Phyllodactylidae) from Isla de Guanaja in the Honduran Bay Islands. Zootaxa 3694: 051-058. Request from author
  • McCranie JR, Hedges SB. 2013. Two additional new species of Sphaerodactylus (Reptilia, Squamata, Gekkonoidea, Sphaerodactylidae) from the Honduran Bay Islands. Zootaxa 3694: 040-050. Request from author
  • Teeling EC, Hedges SB. 2013. Making the impossible possible: rooting the tree of placental mammals. Mol. Biol. Evol. 30: 1999-2000. Advance Access E-print
  • Hedges SB. 2013. Revision shock in taxonomy. Zootaxa 3681: 297-298. Request from author
  • Reynolds RG, Niemiller ML, Hedges SB, Dornburg A, Puente-Rolon AR, Revell LJ. 2013. Molecular phylogeny and historical biogeography of West Indian boid snakes (Chilabothrus). Mol. Phylogenet. Evol. 68: 461-470. E-print
  • Hedges SB. 2013. Caribmap: a mobile-friendly web resource for Caribbean maps that facilitates quantitative historical analysis. Journal of Map and Geography Libraries 9(3): 350-360. Request from author
  • Barrio-Amoros CL, Heinicke MP, Hedges SB. 2013. A new tuberculated Pristimantis (Anura, Terrarana, Strabomantidae) from the Venezuelan Andes, redescription of Pristimantis pleurostriatus, and variation within Pristimantis vanadisae. Zootaxa 3647: 43-62. Request from author
  • Colston TJ, Grazziotin FG, Shepard DB, Vitt LJ, Colli GR, Henderson RW, Hedges SB, Bonatto S, Zaher H, Noonan BP, Burbrink FT. 2013. Molecular systematics and historical biogeography of tree boas (Corallus spp.).Mol. Phylogenet. Evol. 66: 953-959. E-print
  • Kingdon JA, Seiffert ER, Hedges SB, Rathbun G. 2013. Supercohort Afrotheria. Pp. 143-146 in Kingdon JA, Happold D, Butynski T, Hoffmann M, Happold M, Kalina J (Eds.), Mammals of Africa, Volume 1: Introductory chapters and Afrotheria. Bloomsbury Publishing, London. Published 28 February 2013. E-print Purchase book
  • Hedges SB. 2013. Wormholes record species history in space and time. Biology Letters 9 (1), 20120926. PODCAST Request from author
  • Böhm M, et al. 2013. The conservation status of the world's reptiles.Biological Conservation 157: 372-385. Published online 22 December 2012, in print January 2013. E-print

2012

  • Schmid M, Steinlein C, Bogart JP, Feichtinger W, Haaf T, Nanda I, del Pino E, Duellman WE, Hedges SB. 2012. The hemiphractid frogs: phylogeny, embryology, life history, and cytogenetics. Cytogenomics and Genome Research 138: 69-384. Published online, December 2012. Bound as a book (Schmid M, Bogart JP, Hedges SB, Eds). Title page and contents (no reprints available). Order Book
  • Barrio-Amoros CL, Guayasamin JM, Hedges SB. 2012. A new minute Andean Pristimantis (Anura: Strabomantidae) from Venezuela. Phyllomedusa 11: 83-93. E-print
  • Marin J, Donnellan SC, Hedges SB, Doughty P, Hutchinson MN, Cruaud C, Vidal N. 2013. Tracing the history and biogeography of the Australian blindsnake radiation. J. Biogeogr. 40: 928-937, published online 23 November 2012. Request from author
  • McCranie JR, Hedges, SB. 2012. Two new species of geckos from Honduras and resurrection of Sphaerodactylus continentalis Werner from the synonymy of Sphaerodactylus millepunctatus Hallowell (Reptilia, Squamata, Gekkonoidea, Sphaerodactylidae). Zootaxa 3492: 65-76. Request from author
  • Hedges, SB. 2012. Amniote phylogeny and the position of turtles. BMC Biology 10 (64):1-2. E-print
  • Vidal N, Marin J, Sassi J, Battistuzzi FU, Donnellan S, Fitch AJ, Fry BG, Vonk FJ, Rodriguez RC, Couloux A, Hedges SB. 2012. Molecular evidence for an Asian origin of monitor lizards followed by Tertiary dispersals to Africa and Australasia. Biology Letters doi:10.1098/rsbl.2012.0460. E-print
  • Hedges, SB and CE Conn. 2012. A new skink fauna from Caribbean islands (Squamata, Mabuyidae, Mabuyinae). Zootaxa 3288: 1-244. Request from author
  • Díaz, LM, SB Hedges, M Schmid. 2012. A new cryptic species of the genus Eleutherodactylus (Amphibia: Anura: Eleutherodactylidae) from Cuba. Zootaxa 3220: 44-60. Request from author

2011

  • Hedges, SB. 2011. The type species of the threadsnake genus Tricheilostoma Jan revisited (Squamata, Leptotyphlopidae). Zootaxa 3027: 63-64. E-print
  • Kumar S, Hedges, SB. 2011. TimeTree2: species divergence times on the iPhone. Bioinformatics doi:10.1093/bioinformatics/brt315, p. 1-2. E-print
  • Hedges, SB. 2011. On the use of high-level taxonomic names. Zootaxa 2867:67-68. E-print
  • Hedges, SB and LM Díaz. 2011. The conservation status of amphibians in the West Indies. Pp. 31–47 in A. Hailey, B. Wilson, and J. Horrocks (Eds.), Conservation of Caribbean Island Herpetofaunas Volume 1: Conservation Biology and the Wider Caribbean. Brill, Leiden, The Netherlands. E-print
  • Hedges, SB. 2011. An overview of the evolution and conservation of West Indian amphibians and reptiles. Pp. 19–30 in A. Hailey, B. Wilson, and J. Horrocks (Eds.), Conservation of Caribbean Island Herpetofaunas Volume 1: Conservation Biology and the Wider Caribbean. Brill, Leiden, The Netherlands. (reprinted from Applied Herpetology 3:281-292 (2006)). E-print
  • Heinicke MP, Diaz LM, Hedges SB. 2011. Origin of invasive Florida frogs traced to Cuba. Biology Letters, published 26 January 2011, doi 10.1098/rsbl.2010.1131. E-print

2010

  • Schmid M, Steinlein C, Bogart JP, Feichtinger W, Leon P, La Marca E, Diaz LM, Sanz A, Chen S-H, and Hedges SB. 2010. The chromosomes of terraranan frogs: insights into vertebrate cytogenetics. Cytogenet Genome Res 2010; 130–131:1–568. Purchase this book
  • Hoffman, M. et al. 2010. The impact of conservation on the status of the World's vertebrates. Science 330: 1503-1509. E-print
  • Hedges, S.B. 2010. Molecular clocks, flotsam, and Caribbean islands. Pp. 353–354 in Cox, C. B. & P. D. Moore, Biogeography: an ecological and evolutionary Approach. 8th Edition (John Wiley & Sons, Inc., Hoboken, New Jersey). E-print
  • Battistuzzi FU, Filipski A, Hedges SB, & Kumar S. 2010. Performance of relaxed clock methods in estimating evolutionary divergence times and their credibility intervals. Mol. Biol. Evol. 27: 1289-1300. E-print
  • Vidal, N, Marin J, Morini M, Donnellan S, Branch WR, Thomas R, Vences M, Wynn A, Cruaud C, & Hedges SB. 2010. Blindsnake evolutionary tree reveals long history on Gondwana. Biology Letters 6:558-561. E-print Supp

2009

  • Hedges, S. B. 2009. Terrestrial-breeding frogs (Strabomantidae) in Peru. Phyllomedusa 8:147–148. E-print
  • Adalsteinsson, S.A., Branch, W.R., Trape, S., Vitt, L.J., & Hedges, S.B. 2009. Molecular phylogeny, classification, and biogeography of snakes of the Family Leptotyphlopidae (Reptilia, Squamata). Zootaxa 2244: 1-50. Request from author
  • Heinicke, M.P., Duellman, W. E., Trueb, L., Means, D. B., MacCulloch, R. D., Hedges, S. B. 2009. A new frog family (Anura: Terrarana) from South America and an expanded direct-developing clade revealed by molecular phylogeny. Zootaxa 2211: 1-35. Request from author
  • Vidal, N., Rage, J.-C., Couloux, A., Hedges, S. B. 2009. Snakes (Serpentes). Pp. 390-397 in The Timetree of Life, S. B. Hedges and S. Kumar, Eds. (Oxford University Press, New York, 2009). E-print Supplementary Information
  • Hedges, S. B., Vidal, N. 2009. Lizards, snakes, and amphisbaenians (Squamata). Pp. 383-389 in The Timetree of Life, S. B. Hedges and S. Kumar, Eds. (Oxford University Press, New York, 2009). E-print
  • Heinicke, M. P., Sander, J. M., Hedges, S. B. 2009. Lungfishes (Dipnoi). Pp. 348-350 in The Timetree of Life, S. B. Hedges and S. Kumar, Eds. (Oxford University Press, New York, 2009). E-print
  • Heinicke, M. P., Naylor, J. P., Hedges, S. B. 2009. Cartilaginous fishes (Chondrichthyes). Pp. 320-327 in The Timetree of Life, S. B. Hedges and S. Kumar, Eds. (Oxford University Press, New York, 2009). E-print
  • Hedges, S. B. 2009. Vertebrates (Vertebrata). Pp. 309-314 in The Timetree of Life, S. B. Hedges and S. Kumar, Eds. (Oxford University Press, New York, 2009). E-print
  • Bhattacharya, D., Yoon, H.-S., Hedges, S. B., Hackett, J. D. 2009. Eukaryotes (Eukaryota). Pp. 116-120 in The Timetree of Life, S. B. Hedges and S. Kumar, Eds. (Oxford University Press, New York, 2009). E-print
  • Battistuzzi, F. U., Hedges, S. B. 2009. Eubacteria. Pp. 106-115 in The Timetree of Life, S. B. Hedges and S. Kumar, Eds. (Oxford University Press, New York, 2009). E-print
  • Battistuzzi, F. U., Hedges, S. B. 2009. Archaebacteria. Pp. 101-105 in The Timetree of Life, S. B. Hedges and S. Kumar, Eds. (Oxford University Press, New York, 2009). E-print
  • Hedges, S. B. 2009. Life. Pp. 89-98 in The Timetree of Life , S. B. Hedges and S. Kumar, Eds. (Oxford University Press, New York, 2009). E-print
  • Hedges, S. B., Kumar, S. 2009. Discovering the timetree of life. Pp. 3-18 in The Timetree of Life, S. B. Hedges and S. Kumar, Eds. (Oxford University Press, New York, 2009). E-print
  • Hedges, S. B., Kumar, S. 2009. Preface. Pp. xi-xiii in The Timetree of Life, S. B. Hedges and S. Kumar, Eds. (Oxford University Press, New York, 2009). E-print
  • Hedges, S. B and S. Kumar (Eds.). 2009. The Timetree of Life. Oxford University Press. 81 Chapters, 551 p. ISBN 9780199535033. Order The Timetree of Life Book Cover
  • Hedges, S. B., A. Couloux, and N. Vidal. 2009. Molecular phylogeny, classification, and biogeography of West Indian racer snakes of the Tribe Alsophiini (Squamata, Dipsadidae, Xenodontinae). Zootaxa 2067: 1-28. E-print
  • Vidal, N. and S. B. Hedges. 2009. The molecular evolutionary tree of lizards, snakes, and amphisbaenians. CR Biologies 332:129-139. E-print
  • Dίaz, L. M. and S. B. Hedges. 2009. First record of the genus Aristelliger (Squamata: Sphaerodactylidae) in Cuba, with description of a new species. Zootaxa 2028:31-40. Request from author
  • Battistuzzi, F.U., and S.B. Hedges.2009. A major clade of prokaryotes with ancient adaptations to life on land. Mol. Biol. Evol. 26:335–344. E-print Supplementary data

2008

  • Vidal, N., W.R. Branch, O.S.G. Pauwels, S.B. Hedges, D.G. Broadley, M. Wink, C. Cruaud, U. Joger & Z.T. Nagy. 2008. Dissecting the major African snake radiation: a molecular phylogeny of the Lamprophiidae Fitzinger (Serpentes, Caenophidia). Zootaxa 1945:51–66. Request from author
  • Hedges, S. B. 2008. Threatened amphibians of the West Indies. Pp. 104–105 in Stuart, S.N., Hoffmann, M., Chanson, J.S., Cox, N.A., Berridge, R., Ramani, P. and Young B.E.(Ed.), Threatened Amphibians of the World. Lynx Ediciones, with IUCN - The World Conservation Union, Conservation International, and NatureServe, Barcelona.
  • Bolaños,F., Castro,F., Cortez,C., De la Riva,I., Grant,T., Hedges,S. B., Heyer,W. R., Ibáñez,R., La Marca,E., Lavilla,E., Silvano,D.L., Lötters,S., Parra Olea,G., Reichle,S., Reynolds,R., Rodríguez,L., Santos Barrera,G., Scott,N. J., Ubeda,C., Veloso,A., Wilkinson,M. and Young, B. 2008. Amphibians of the Neotropical Realm. Pp. 92–99 in Stuart, S.N., Hoffmann, M., Chanson, J.S., Cox, N.A., Berridge, R., Ramani, P. and Young B.E. (Ed.), Threatened Amphibians of the World. Lynx Ediciones, with IUCN - The World Conservation Union, Conservation International, and NatureServe, Barcelona.
  • Hedges, S. B. 2008. Dating old maps with the print clock. The Portolan 72: 25-33. E-print
  • Hedges, S. B. 2008. At the lower limit of size in snakes: two new species of threadsnakes (Squamata: Leptotyphlopidae: Leptotyphlops) from the Lesser Antilles. Zootaxa 1841:1–30. E-print
  • Hedges, S. B., W. E. Duellman, and M. P. Heinicke. 2008. A replacement name for Isodactylus Hedges, Duellman, and Heinicke, 2008. Zootaxa 1795:67-68. Request from author
  • Dίaz, L. M. and S. B. Hedges. 2008. A new gecko of the genus Tarentola (Sauria: Gekkonidae) from eastern Cuba. Zootaxa 1743:43–52. Request from author
  • Hedges, S. B., W. E. Duellman, and M. P. Heinicke. 2008. New World direct-developing frogs (Anura: Terrarana): molecular phylogeny, classification, biogeography, and conservation. Zootaxa 1737:1–182. Request from author
  • Hedges, S. B. 2008. Image analysis of Renaissance copperplate prints. Proc. SPIE 6810: 681009. 20 pages. E-print
  • Duellman, W. E. and S. B. Hedges. 2008. Two new minute species of Phrynopus (Lissamphibia: Anura) from the Cordillera Oriental in Peru. Zootaxa 1675:59–66. Request from author
  • Vidal, N., A. Azvolinsky, C. Cruaud, and S. B. Hedges. 2008. Origin of tropical American burrowing reptiles by transatlantic rafting. Biology Letters 4:115-118. Published online 11 December 2007 and in print 7 January 2008. E-print

2007

  • Duellman, W. E. and S. B. Hedges. 2007. Three new species of Pristimantis (Lissamphibia: Anura) from Montane Forests of the Cordillera Yanachaga in Central Peru. Phyllomedusa 6:119–135. E-print
  • Lehr, E., J. B. Pramuk, S. B. Hedges, and J. H. Córdova. 2007. A new species of arboreal Rhinella (Anura: Bufonidae) from Yanachaga-Chemillén National Park in central Peru. Zootaxa 1662:1–14. Request from author
  • Hedges, S. B. 2007. The amphibians and reptiles of Hispaniola and their conservation. Pp. 18–25 in Fernandez, E. M. (Ed.), Hispaniola: a photographic journey through island biodiversity. Harvard University Press, Cambridge, Massachusetts. E-print
  • Heinicke, M. P., W. E. Duellman, and S. B. Hedges. 2007. Major Caribbean and Central American frog faunas originated by ancient oceanic dispersal. Proc. Natl. Acad. Sci. (USA) 104:10092-10097. E-print SI text and tables
  • Vidal, N., A.-S. Delmas, and S. B. Hedges. 2007. The higher-level relationships of alethinophidian snakes inferred from seven nuclear and mitochondrial genes. Pp. 27-33 in R. W. Henderson and R. Powell (Eds.), Biology of the boas and pythons. Eagle Mountain Publishing, Eagle Mountain, Utah. E-print
  • Weiss, A. J., and S. B. Hedges. 2007. Molecular phylogeny and biogeography of the Antillean geckos Phyllodactylus wirshingi, Tarentola americana, and Hemidactylus haitianus (Reptilia, Squamata). Mol. Phylo. Evol. 45:409-416. E-print
  • Hedges, S. B., and M. P. Heinicke. 2007. Molecular phylogeny and biogeography of West Indian frogs of the genus Leptodactylus. Mol. Phylo. Evol. 44:308-314. E-print
  • Vidal, N., A. S. Delmas, P. David, C. Cruaud, A. Couloux, and S. B. Hedges. 2007. The phylogeny and classification of caenophidian snakes inferred from seven nuclear protein-coding genes. CR Biologies 330:182-187. E-print
  • Thomas, R. and S. B. Hedges. 2007. Eleven new species of snakes of the genus Typhlops (Serpentes: Typhlopidae) from Hispaniola and Cuba. Zootaxa 1400:1-26. Request from author

2006

  • Thomas, R. and S. B. Hedges. 2006. Two new species of Amphisbaena (Reptilia: Squamata: Amphisbaenidae) from the Tiburon Peninsula of Haiti. Caribb. J. Sci. 42:208-219. E-print (published in early 2007).
  • Hedges, S. B., F. U. Battistuzzi, and J. E. Blair. 2006. Molecular timescale of evolution in the Proterozoic. Pp. 199-229 in S. Xiao and A. J. Kaufman (Eds.), Neoproterozoic Geobiology and Paleobiology, Springer, New York. E-print
  • Hedges, S. Blair. 2006. An overview of the evolution and conservation of West Indian amphibians and reptiles. Applied Herpetology 3:281-292. E-print
  • Hedges, S. B., J. Dudley, and S. Kumar. 2006. TimeTree: a public knowledge-base of divergence times among organisms. Bioinformatics 22:2971-2972. E-print
  • Dominguez, M., L. V. Moreno, and S. B. Hedges. 2006. A new snake of the genus Tropidophis (Tropidophiidae) from the Guanahacabibes Peninsula of western Cuba. Amphibia-Reptilia 27:427-432. E-print
  • Hedges, S. B. 2006. Paleogeography of the Antilles and the origin of West Indian Terrestrial Vertebrates. Ann. Missouri Bot. Gard. 93:231-244. E-print
  • Hedges, S. B., S. Kumar, and M. van Tuinen. 2006. Constraining fossil calibrations for molecular clocks. Bioessays 28:770-771. E-print
  • Hedges, S. B. 2006. A method for dating early books and prints using image analysis. Proc. R. Soc. Lond. A: Mathematical, Physical, and Engineering Sciences 462:3555-3573. E-print Appendix
  • Hedges, S. B. 2006.Singularities: landmarks on the pathways of life. Quart. Rev. Biol. 81:164. (book review). E-print
  • van Rheede, T., T. Bastiaans, D. N. Boone, S. B. Hedges, W. W. de Jong, and O. Madsen. 2006. The platypus is in its place: nuclear genes and indels confirm the sister group relation of monotremes and therians. Mol. Biol. Evol. 23:587-597. E-print
  • Fry, B.G., N. Vidal, J. A. Norman, F. J. Vonk, H. Scheib, R. Ramjan, S. Kuruppu, K. Fung, S. B. Hedges, M. K. Richardson, W. C. Hodgson, V. Ignjatovic, R. Summerhayes, and E. Kochva. 2006. Early evolution of the venom system in lizards and snakes. Nature 439:584-588(online 17 November 2005). E-print

2005

  • Kumar, S. and S. B. Hedges. 2005. Pushing back the expansion of introns in animal genomes. Cell 123:1182-1184. E-print
  • Kumar, S., A. Filipski, V. Swarna, A. Walker & S. B. Hedges. 2005. Placing confidence limits on the molecular age of the human-chimpanzee divergence. Proc. Natl. Acad. Sci. 102:18842-18847. E-print suppl info
  • Vidal, N., and S. B. Hedges. 2005. The phylogeny of squamate reptiles (lizards, snakes, and amphisbaenians) inferred from nine nuclear protein-coding genes. C. R. Biologies 328:1000-1008. E-print
  • Blair, J. E., and S. B. Hedges. 2005. Molecular phylogeny and divergence times of deuterostome animals. Mol. Bio. Evol. 22:2275-2284. E-print
  • Vidal, N., R. W. Henderson, A.-S. Delmas, and S. B. Hedges. 2005. A phylogenetic study of the emerald treeboa (Corallus caninus). J. Herpetology 39:500-503. E-print
  • Duellman, W. E., and S. B. Hedges. 2005. Eleutherodactyline frogs from the Cordillera Yanachaga in Central Peru. Copeia 2005:526-538. E-print
  • Smith, M. L., S. B. Hedges, W. Buck, A. Hemphill, S. Inchaustegui, M. Ivie, D. Martina, M. Maunder, and J. F. Ortega. 2005. Caribbean Islands. Pp 112-118 in R. A. Mittermeier, P. R. Gil, M. Hoffman, J. Pilgrim, T. Brooks, C. G. Mittermeier, J. Lamoreux, and G. A. B. da Fonseca (eds.), Hotspots revisited: Earth's biologically richest and most endangered terrestrial ecoregions. Mexico City: CEMEX. E-print
  • M. Schmid, I. Nanda, H. Hoehn, M. Schartl, T. Haaf, J.-M. Buerstedde, H. Arakawa, R. B. Caldwell, S. Weigend, D. W. Burt, J. Smith, D. K. Griffin, J. S. Masabanda, M. A. M. Groenen, R. P. M. A. Crooijmans, A. Vignal, V. Fillon, M. Morisson, F. Pitel, M. Vignoles, A. Garrigues, J. Gellin, A. V. Rodionov, S. A. Galkina, N. A. Lukina, G. Ben-Ari, S. Blum, J. Hillel, T. Twito, U. Lavi, L. David, M. W. Feldman, M. E. Delany, C.A. Conley, V. M. Fowler, S. B. Hedges, R. Godbout, S. Katyal, C. Smith, Q. Hudson, A. Sinclair and S. Mizunou. 2005. Second report on chicken genes and chromosomes 2005. Cytogenet. Genome Res. 109: 415-479. E-print
  • Blair, J. E., P. Shah, and S. B. Hedges. 2005. Evolutionary sequence analysis of complete eukaryote genomes. BMC Bioinformatics 4(53):1-10. E-print
  • Nicholson, K. E., R. E. Glor, J. J. Kolbe, A. Larson, S. B. Hedges, and J. B. Losos. 2005. Mainland colonization by island lizards. J. Biogeogr. 32:1-10. E-print
  • Blair, J. E., and S. B. Hedges. 2005. Molecular clocks do not support the Cambrian explosion. Mol. Biol. Evol. 22:387-390. E-print

2004

  • Young, B. E., and S. B. Hedges. 2004. Haiti: a megadiverse Caribbean country. P. 20 in B. E. Young, S. N. Stuart, J. S. Chanson, N. A. Cox, and T. M. Boucher, Disappearing Jewels: the Status of New World Amphibians. NatureServe, Arlington, Virginia.
  • Rabosky, D. L., K. P. Aplin, S. C. Donnellan, and S. B. Hedges. 2004. Molecular phylogeny of blindsnakes (Ramphotyphlops) from Western Australia and resurrection of Ramphotyphlops bicolor (Peters, 1857). Australian J. Zool. 52:531-548. E-print
  • Battistuzzi, F. U., A. Feijão, and S. B. Hedges. 2004. A genomic timescale of prokaryote evolution: insights into the origin of methanogenesis, phototrophy, and the colonization of land. BMC Evol. Biol. 4:44 (14 pages). E-print
  • Hedges, S. B. and S. Kumar. 2004. Precision of molecular time estimates. Trends in Genetics 20:242-247. E-print
  • Vidal, N. and S. B. Hedges. 2004. Molecular evidence for a terrestrial origin of snakes. Proc. R. Soc. Lond. B (Suppl.) 271:S226-S229. E-print
  • Hedges, S. B., J. E. Blair, M. L. Venturi, and J. L. Shoe. 2004. A molecular timescale of eukaryote evolution and the rise of complex multicellular life. BMC Evol. Biol. 4:2 (9 pages). E-print
  • Pisani, D., L. L. Poling, M. Lyons-Weiler, and S. B. Hedges. 2004. The colonization of land by animals: molecular phylogeny and divergence times among arthropods. BMC Biology 2:1 (10 pages). E-print
  • van Tuinen, M., and S. B. Hedges. 2004. The effect of external and internal fossil calibrations on the avian evolutionary timescale. Journal of Paleontology 78:45-50. E-print

2003

  • Hower, L. M. and S. B. Hedges. 2003. Molecular Phylogeny and Biogeography of West Indian Teiid Lizards of the Genus Ameiva. Caribb. J. Sci. 39:298-306. E-print
  • Hedges, S. B. 2003. Molecular clocks and a biological trigger for the Neoproterozoic snowball Earths and Cambrian explosion. Pp 27-40 in P. Donoghue and P. Smith, Eds., Telling evolutionary time: molecular clocks and the fossil record. CRC Press, London. E-print
  • Hedges, S. B. 2003. The coelacanth of frogs. Nature 425:669-670. E-print
  • Des Marais, D. J., L. J. Allamandola, S. A. Benner, A. P. Boss, J. R. Cronin, D. Deamer, P. G. Falkowski, J. D. Farmer, S. B. Hedges, B. M. Jakosky, A. H. Knoll, D. R. Liskowsky, V. J. Meadows, M. A. Meyer, C. B. Pilcher, K. H. Nealson, A. M. Spormann, J. D. Trent, W. W. Turner, N. J. Woolf, and H. W. Yorke. 2003. The NASA Astrobiology Roadmap. Astrobiology 3:219-235. E-print
  • Hedges, S. B. 2003. The frog detectives of Pico Turquino. Pp. 51-57 in R. W. Henderson and R. Powell (eds.), Islands and the Sea: Essays on the Herpetological Exploration in the West Indies. Society for the Study of Amphibians and Reptiles, Ithaca (New York).
  • Diaz, L. M., A. Cadiz, and S. B. Hedges. 2003. A new grass frog from pine forests of western Cuba, and description of acoustic and pattern variation in Eleutherodactylus varleyi (Amphibia: Leptodactylidae). Caribb. J. Sci. 39:176-188. E-print
  • Hedges, S. B. and P. Shah. 2003. Comparison of mode estimation methods and application in molecular clock analysis. BMC Bioinformatics 4:31. E-print
  • Hedges, S. B., and S. Kumar. 2003. Genomic clocks and evolutionary timescales. Trends in Genetics 19:200-206. E-print

2002

  • Werner, T., S. Weissman, S. B. Hedges, and K. Gardiner. 2002. Report on the twelfth international workshop on the identification of transcribed sequences: functional, expression, and evolutionary analysis. Cytogenet. Genome Res. 98:109-117.
  • Hedges, S. B. 2002. Early life (book review). Quart. Rev. Biol. 77:445. E-print
  • Hedges, S. B. 2002. Morphological variation and the definition of species in the snake genus Tropidophis (Tropidophiidae). Bull. Nat. Hist. Mus. (London), Zool. Ser. 68:83-90. E-print
  • Vidal, N., and S. B. Hedges. 2002. Higher-level relationships of caenophidian snakes inferred from four nuclear and mitochondrial genes. C. R. Biologies 325:987-995. E-print
  • Vidal, N., and S. B. Hedges. 2002. Higher-level relationships of snakes inferred from four nuclear and mitochondrial genes. C. R. Biologies 325:977-985. E-print
  • Hedges, S. B. 2002. The origin and evolution of model organisms. Nature Reviews Genetics 3:838-849. E-print
  • Hedges, S. B., and S. Kumar. 2002. Vertebrate genomes compared. Science 297:1283-1285. E-print
  • Highton, R., S. B. Hedges, C. A. Hass, and H. G. Dowling. 2002. Snake relationships revealed by slowly-evolving proteins: further analysis and a reply. Herpetologica 58: 270-275. E-print
  • Hedges, S. B., and O. H. Garrido. 2002. A new snake of the genus Tropidophis (Tropidophiidae) from Eastern Cuba. J. Herpetology 36:157-161. E-print
  • Blair, J. E., K. Ikeo, T. Gojobori, and S. B. Hedges. 2002. The evolutionary position of nematodes. BMC Evolutionary Biology 2:7 (7 pages). E-print

2001

  • Stauffer, R. L., A. Walker, O. Ryder, M. Lyons-Weiler, and S. B. Hedges. 2001. Human and ape molecular clocks and constraints on paleontological hypotheses. J. Heredity 92:469-474. E-print
  • Hedges, S. B., O. H. Garrido, and L. M. Diaz. 2001. A new banded snake of the genus Tropidophis (Tropidophiidae) from North-Central Cuba. J. Herpetology 35:615-617. E-print
  • Hedges, S. B., and R. Thomas. 2001. At the lower size limit in amniotes: a new diminutive lizard from the West Indies. Caribbean J. Sci. 37:168-173. E-print
  • Garrido, O. H., and S. B. Hedges. 2001. A new anole from the northern slope of the Sierra Maestra in eastern Cuba. J. Herpetology 35: 378-383. E-print
  • Hedges, S. B., H. Chen, S. Kumar, D. Y.-C. Wang, A. S. Thompson, and H. Watanabe. 2001. A genomic timescale for the origin of eukaryotes. BMC Evolutionary Biology 1:4 (10 pages). E-print
  • Pramuk, J. B., C. A. Hass, and S. B. Hedges. 2001. Molecular phylogeny and biogeography of West Indian toads (Anura: Bufonidae). Mol. Phylogenet. Evol. 20:294-301. E-print
  • Heckman, D. S., D. M. Geiser, B. R. Eidell, R. L. Stauffer, N. L. Kardos, and S. B. Hedges. 2001. Molecular evidence for the early colonization of land by fungi and plants. Science293:1129-1133. E-print
  • van Tuinen, M., D. B. Butvill, J. A. W. Kirsch, and S. B. Hedges. 2001. Convergence and divergence in the evolution of aquatic birds. Proc. Roy. Soc. London B268: 1345-1350. E-print
  • Hass, C. A., L. R. Maxson, and S. B. Hedges. 2001. Relationships and divergence times of West Indian amphibians and reptiles: insights from albumin immunology. In C. A. Woods and F. E. Sergile (eds.), Biogeography of the West Indies: patterns and perspectives. CRC Press, Boca Raton, Florida. E-print
  • Hedges, S. B. 2001. Biogeography of the West Indies: an overview. In C. A. Woods and F. E. Sergile (eds.), Biogeography of the West Indies: patterns and perspectives. CRC Press, Boca Raton, Florida. E-print
  • Diaz, L. M., A. E. Estrada, and S. B. Hedges. 2001. A new riparial frog of the genus Eleutherodactylus (Anura: Leptodactylidae) from Eastern Cuba. Caribbean J. Sci. 37: 63-71. E-print
  • Hedges, S. B. 2001. Molecular evidence for the early history of living vertebrates. Pp. 119-134 in P. E. Ahlberg (Ed.) Major events in early vertebrate evolution: palaeontology, phylogeny, genetics and development. Taylor and Francis, London. E-print
  • van Tuinen, M., and S. B. Hedges. 2001. Calibration of avian molecular clocks. Mol. Biol. Evol. 18:206-213. E-print
  • Hedges, S. B. 2001. Afrotheria: plate tectonics meets genomics. Proc. Nat. Acad. Sci. (USA) 98:1-2. E-print

2000

  • Tracy, M. R. and S. B. Hedges. 2000. Evolutionary history of the enolase gene family. Gene 259:129-138. E-print
  • Hedges, S. B. 2000. A start for population genomics. Nature 408:652-653. E-print
  • Garrido, O. H. and S. B. Hedges. 2000. Elevation of Anolis vanidicus rejectus Garrido and Schwartz (Sauria: Iguanidae) to species status and designation of Anolis mimus Schwartz and Thomas as a synonym. Caribb. J. Sci. 36:20-21. E-print
  • Vidal, N., S. G. Kindl, A. Wong, and S. B. Hedges. 2000. Phylogenetic relationships of xenodontine snakes inferred from 12S and 16S ribosomal RNA sequences. Mol. Phylogenet. Evol. 14:389-402. E-print
  • van Tuinen, M., C. G. Sibley, and S. B. Hedges. 2000. The early history of modern birds inferred from DNA sequences of nuclear and mitochondrial ribosomal genes. Mol. Biol. Evol. 17:451-457. E-print
  • Hedges, S. B., and M. H. Schweitzer. 1999. Molecular paleontology. Pp. 752-754 in R. Singer (ed.) Encyclopedia of Paleontology. Fitzroy-Dearborn Publishers, Chicago. E-print
  • Hedges, S. B. 1999. Caught in the act. Nature 402:459. (Book review). E-print
  • O'Brien, S. J., J. F. Eisenberg, M. Miyamoto, S. B. Hedges, S. Kumar, and D. E. Wilson. 1999. Comparative genomics: mammalian radiations. Genome Maps 10 (phylogenetic tree). Science 286:463-478. E-print
  • Hedges, S. B., and S. Kumar. 1999. Divergence times of eutherian mammals. Science 285:2031. E-print
  • Hedges, S. B., and O. H. Garrido. 1999. A new snake of the genus Tropidophis (Tropidophiidae) from central Cuba. J. Herpetology 33:436-441. E-print
  • Hedges, S. B. 1999. Distribution patterns of amphibians in the West Indies. Pp. 211-254 In, Duellman, W. E., Regional Patterns of Amphibian Distribution: A Global Perspective. Johns Hopkins University Press. E-print
  • Hedges, S. B., A. R. Estrada, and L. M. Diaz. 1999. A new snake (Tropidophis) from western Cuba. Copeia 1999:376-381. E-print
  • Hedges, S. B., and L. L. Poling. 1999. A molecular phylogeny of reptiles. Science 283:998-1001. E-print
  • Wang, D. Y.-C., S. Kumar, and S. B. Hedges. 1999. Divergence time estimates for the early history of animal phyla and the origin of plants, animals, and fungi. Proc. Roy. Soc. London B 266:163-171. E-print
  • Johnson, K., and S. B. Hedges. 1998. Three new species of Calisto from southwestern Haiti (Lepidoptera: Nymphalidae: Satyrinae). Tropical Lepidoptera 9:45-53. E-print
  • Hedges, S. B., and R. Powell. 1998. Eleutherodactylus thorectes. Cat. Amer. Amph. Rept. 667:1-2. E-print
  • Estrada, A. R., and S. B. Hedges. 1998. Sistematica de las ranas riberenas de Cuba (Leptodactylidae: Eleutherodactylus). Carib. J. Sci. 34:218-230. E-print
  • Hedges, S. Blair. 1998. Nature Debates: Molecules and the fossil record. Nature (www), November 1998. (Invited Essay). E-print
  • Thomas, R. and S. B. Hedges. 1998. A new gecko from the Sierra de Neiba of Hispaniola. Herpetologica 54:333-336. E-print
  • Stanhope, M. J., V. G. Waddell, O. Madsen, W. de Jong, S. B. Hedges, G. Cleven, D. Kao, and M. S. Springer. 1998. Molecular evidence for multiple origins of Insectivora and for a new order of endemic African insectivore mammals. Proc. Nat. Acad. Sci. (USA) 95:9967-9972. E-print
  • Feller, A. E. and S. B. Hedges. 1998. Molecular evidence for the early history of living amphibians. Mol. Phylogenet. Evol. 9:509-516. E-print
  • Schubart, C., R. Diesel, and S. B. Hedges. 1998. Rapid evolution to terrestrial life in Jamaican crabs. Nature 393:363-365. E-print
  • Kumar, S., and S. B. Hedges. 1998. A molecular timescale for vertebrate evolution. Nature 392:917-920. E-print
  • van Tuinen, M., C. G. Sibley, and S. B. Hedges. 1998. Phylogeny and biogeography of ratite birds inferred from DNA sequences of the mitochondrial ribosomal genes. Mol. Biol. Evol. 15:370-376. E-print
  • Thomas, R. and S. B. Hedges. 1998. A new amphisbaenian from Cuba. J. Herpetol. 32:92-96. E-print
  • Thomas, R., S. B. Hedges, and O. H. Garrido. 1998. A new gecko (Sphaerodactylus) from the Sierra Maestra of Cuba. J. Herpetol. 32:66-69. E-print
  • Hedges, S. B., and R. Powell. 1998. Eleutherodactylus parapelates. Cat. Amer. Amph. Rept. 649:1-2. E-print
  • Cunningham, C. A., S. B. Hedges, and R. Powell. 1998. Eleutherodactylus chlorophenax. Cat. Amer. Amph. Rept 648:1-2. E-print
  • Hedges, S. B., and R. Powell. 1998. Eleutherodactylus amadeus. Cat. Amer. Amph. Rept. 647:1-2. E-print
  • Hedges, S. B. 1998. Avian Molecular Evolution and Systematics. Amer. Zool. 37:262-263. (Book review). E-print
  • Thomas, R., and S. B. Hedges. 1998. New anguid lizard from Cuba. Copeia 1998:97-103. E-print
  • Hedges, S. B., and L. R. Maxson. 1997. Complementary uses of molecules and morphology. Mol. Phylogenet. Evol. 8:445. E-print
  • Estrada, A. R., and S. B. Hedges. 1997. A new frog of the genus Eleutherodactylus from the Meseta de Cabo Cruz, eastern Cuba (Anura, Leptodactylidae). Carib. J. Sci. 33:227-232. E-print
  • Estrada, A. R., and S. B. Hedges. 1997. Nueva especie de Eleutherodactylus (Anura: Leptodactylidae) del Macizo de Sagua-Baracoa, región oriental de Cuba. Carib. J. Sci. 33:222-226. E-print
  • Estrada, A. R., and S. B. Hedges. 1997. A new species of frog from the Sierra Maestra, eastern Cuba (Leptodactylidae, Eleutherodactylus). J. Herpetol. 31:364-368. E-print
  • Hedges, S. B. 1997. Amphibians and Reptiles of the Yucatán Peninsula. Quart. Rev. Biol. 72:213. (Book review). E-print
  • Hedges, S. B. 1997. Molecular zoology. Biochem. Syst. Ecol. 25:187. (Book review). E-print
  • Hedges, S. B. 1996. More on Caribbean zoogeography. Herpetologica 52:622-624. E-print
  • Hedges, S. B. 1996. Historical biogeography of West Indian vertebrates. Ann. Rev. Ecol. Syst. 27:163-196. E-print
  • Estrada, A. R., and S. B. Hedges. 1996. At the lower size limit in tetrapods: a new diminutive frog from Cuba (Leptodactylidae: Eleutherodactylus). Copeia 1996:852-859. E-print
  • Hedges, S. B., and L. R. Maxson. 1996. Molecules and morphology in amniote phylogeny. Mol. Phylogenet. Evol. 6:312-314. E-print
  • Dowling, H. G., C. A. Hass, S. B. Hedges, and R. Highton. 1996. Snake relationships revealed by slow-evolving proteins: a preliminary survey. J. Zool. (London) 240:1-28. E-print
  • Hedges, S. B. 1996. Vicariance and dispersal in Caribbean biogeography. Herpetologica 52:466-473. E-print
  • Estrada, A. R., and S. B. Hedges. 1996. A new frog of the genus Eleutherodactylus from eastern Cuba (Anura, Leptodactylidae). Herpetologica 52:435-439. E-print
  • Hedges, S. B. 1996. The Hispaniolan frog Eleutherodactylus neodreptus Schwartz (Anura: Leptodactylidae) is a synonym of E. audanti Cochran. Caribb. J. Sci. 32:248. E-print
  • Hedges, S. B., P. H. Parker, C. G. Sibley, and S. Kumar. 1996. Continental breakup and the ordinal diversification of birds and mammals. Nature 381:226-229. E-print
  • Hedges, S. B. 1996. The origin of West Indian amphibians and reptiles. Pp 95-128 in R. Powell and R. W. Henderson (eds.). Contributions to West Indian Herpetology: A tribute to Albert Schwartz. Society for the Study of Amphibians and Reptiles, Ithaca, NY. E-print
  • Hedges, S. B., L. Gonzalez, and A. R. Estrada. 1995. Rediscovery of the Cuban frogs Eleutherodactylus cubanus and E. turquinensis (Anura: Leptodactylidae). Caribb. J. Sci. 31:327-332. E-print
  • Hedges, S. B., Simmons, M. D., VanDikj, M. A., Caspers, G., DeJong, W. W., and C. G. Sibley. 1995. Phylogenetic relationships of the Hoatzin, an enigmatic South American bird. Proc. Nat. Acad. Sci. (USA)92:11662-11665. E-print
  • Highton, R. and S. B. Hedges. 1995. Geographic protein variation in Pseudacris brimleyi (Anura: Hylidae) analysis by sequential electrophoresis. J. Herpetol. 29:351-357. E-print
  • Hay, J. M., I. Ruvinsky, S. B. Hedges, and L. R. Maxson. 1995. Phylogenetic relationships of amphibian families inferred from DNA sequences of mitochondrial 12S and 16S ribosomal RNA genes. Mol. Biol. Evol. 12:928-937. E-print
  • Estrada, A. R., and S. B. Hedges. 1995. A new species of Anolis (Sauria: Iguanidae) from eastern Cuba. Caribb. J. Sci. 31:65-72. E-print
  • Hedges, S. B., and M. H. Schweitzer. 1995. Detecting dinosaur DNA. Science 268:1191-1192. E-print
  • Henderson, R. W., and S. B. Hedges. 1995. Origin of West Indian populations of the geographically widespread boa Corallus enydris inferred from mitochondrial DNA sequences. Mol. Phylogenet. Evol. 4:88-92. E-print
  • Heise, P. J., H. G. Dowling, L. R. Maxson, and S. B. Hedges. 1995. Higher-level snake phylogeny inferred from mitochondrial DNA sequences of 12SrRNA and 16SrRNA genes. Mol. Biol. Evol. 12:259-265. E-print
  • Bogart, J. P., and S. B. Hedges. 1995. Rapid chromosome evolution in Jamaican frogs of the genus Eleutherodactylus (Leptodactylidae). J. Zool. (London) 235:9-31. E-print
  • Frye, M. S., and S. B. Hedges. 1995. Monophyly of the order Rodentia inferred from mitochondrial DNA sequences of the genes for 12S rRNA, 16S rRNA, and tRNA-valine. Mol. Biol. Evol. 12:168-176. E-print
  • Hedges, S. B., and K. Johnson. 1994. Calisto tasajera in the Hispaniolan Cordillera Central (Lepidoptera: Nymphalidae: Satyrinae). Trop. Lepidop. 5:93-94. E-print
  • Hedges, S. B., C. A. Hass, and L. R. Maxson. 1994. Towards a biogeography of the Caribbean. Cladistics10:43-55. E-print
  • Hedges, S. B., and C. G. Sibley. 1994. Molecules vs. morphology in avian evolution: the case of the "pelecaniform" birds. Proc. Nat. Acad. Sci. (USA) 91:9861-9865. E-print
  • Hedges, S. B., and R. L. Bezy. 1994. Xantusiid lizards and phylogenetic inference. Mol. Phylogenet. Evol. 3:275-278. E-print
  • Hedges, S. B. 1994. Molecular evidence for the origin of birds. Proc. Natl. Acad. Sci. (U.S.A.) 91:2621-2624. E-print
  • Ruth, B. C., W. A. Dunson, C. L. Rowe, and S. B. Hedges. 1993. Molecular and functional evaluation of the egg mass color polymorphism of the spotted salamander, Ambystoma maculatum. J. Herpetology 27:307-314. E-print
  • Hedges, S. B., and O. H. Garrido. 1993. A new species of gecko (Sphaerodactylus) from central Cuba. J. Herpetology 27:300-306. E-print
  • Thomas, R., and S. B. Hedges. 1993. A new banded Sphaerodactylus from eastern Hispaniola (Squamata: Gekkonidae). Herpetologica 49:350-354. E-print
  • Hedges, S. B., and C. A. Woods. 1993. Caribbean hot spot. Nature 364:375. E-print
  • Hedges, S. B., R. A. Nussbaum, and L. R. Maxson. 1993. Caecilian phylogeny and biogeography inferred from mitochondrial DNA sequences of the 12S rRNA and 16S rRNA genes (Amphibia: Gymnophiona). Herpetol. Monogr. 7:64-76. E-print
  • Hedges, S. B., and L. R. Maxson. 1993. A molecular perspective on Lissamphibian phylogeny. Herpetol. Monogr. 7:27-42. E-print
  • Hedges, S. B. 1993. Global amphibian declines: a perspective from the Caribbean. Biodiversity and Conservation 2:290-303. E-print
  • Hedges, S. B. and R. L. Bezy. 1993. Phylogeny of xantusiid lizards: concern for data and analysis. Mol. Phylogenetics Evol. 2:76-87. E-print
  • Hedges, S. B., C. A. Hass, and L. R. Maxson. 1993. Relations of fish and tetrapods. Nature 363:501-502. E-print
  • Hass, C. A., S. B. Hedges, and L. R. Maxson. 1993. Molecular insights into the relationships and biogeography of West Indian anoline lizards. Biochem. Syst. Ecol. 21:97-114. E-print
  • Hedges, S. B. and A. Schluter. 1992. Eleutherodactylus eurydactylus, a new species of frog from central Amazonian Perú (Anura: Leptodactylidae). Copeia 1992:1002-1006. E-print
  • Thomas, R., and S. B. Hedges. 1992. An unusual new Sphaerodactylus from Hispaniola (Squamata, Gekkonidae). J. Herpetology 26:289-292. E-print
  • Thomas, R., S. B. Hedges, and O. H. Garrido. 1992. Two new species of Sphaerodactylus from eastern Cuba (Sauria, Gekkonidae). Herpetologica 48:358-367. E-print
  • Hedges, S. B., and R. Thomas. 1992. Two new species of Eleutherodactylus from remnant cloud forest in Haiti (Anura, Leptodactylidae). Herpetologica 48:351-358. E-print
  • Hedges, S. B., and O. H. Garrido. 1992. A new species of Tropidophis from Cuba (Serpentes, Tropidophiidae). Copeia1992:820-825. E-print
  • Garrido, O. H., and S. B. Hedges. 1992. Three new grass anoles (Sauria: Iguanidae) from Cuba. Caribb. J. Sci. 28:21-29. E-print
  • Hedges, S. B. 1992. A reconsideration of two montane species of Eleutherodactylus in Hispaniola (Anura, Leptodactylidae). Caribb. J. Sci. 28:11-16. E-print
  • Hedges, S. B., A. R. Estrada, and R. Thomas. 1992. Three new species of Eleutherodactylus from eastern Cuba, with notes on vocalizations of other species (Anura, Leptodactylidae). Herpetol. Monogr. 6:68-83. E-print
  • Hedges, S. B., and O. H. Garrido. 1992. Cuban snakes of the genus Arrhyton: Two new species and a reconsideration of A. redimitum Cope. Herpetologica 48:168-177. E-print
  • Hedges, S. B., and R. Thomas. 1992. A new marsh-dwelling species of Eleutherodactylus from Haiti (Anura:Leptodactylidae). J. Herpetology 26:191-195. E-print
  • Hass, C. A., and S. B. Hedges. 1992. Karyotype of the Cuban lizard Cricosaura typica and its implications for xantusiid phylogeny. Copeia 1992(2):563-565. E-print
  • Hedges, S. B., J. P. Bogart, and L. R. Maxson. 1992. Ancestry of unisexual salamanders (genus Ambystoma). Nature 356:708-710. E-print
  • Hedges, S. B., C. A. Hass, and L. R. Maxson. 1992. Caribbean biogeography: Molecular evidence for dispersal in West Indian terrestrial vertebrates. Proc. Natl. Acad. Sci. (U.S.A.) 89:1909-1913. E-print
  • Hedges, S. B., and L. R. Maxson. 1992. 18S rRNA sequences and amniote phylogeny: reply to Marshall. Mol. Biol. Evol. 9(2):374-377. E-print
  • Hedges, S. B. 1992. The number of replications needed for accurate estimation of the bootstrap p-value in phylogenetic studies. Mol. Biol. Evol. 9(2):366-369. E-print
  • Hedges, S. B., S. Kumar, K. Tamura, and M. Stoneking. 1992. Human origins and analysis of mitochondrial DNA sequences. Science 255:737-739. E-print
  • Hedges, S. B. and R. Thomas. 1991. The importance of systematic research in the conservation of amphibian and reptile populations. Pp. 56-61 in Moreno, J. A. (ed.), Status y distribución de los reptiles y anfibios de la región de Puerto Rico. Departamento de Recursos Naturales de Puerto Rico, Publicación Científica Miscelánea No. 1. E-print
  • Estrada, A. R., and S. B. Hedges. 1991. Nueva especie de Eleutherodactylus (Anura, Leptodactylidae) de la región oriental de Cuba. Caribb. J. Sci. 27:139-145. E-print
  • Schwartz, A., and S. B. Hedges. 1991. An elevational transect of Lepidoptera on Pico Turquino, Cuba. Caribb. J. Sci. 27:130-138. E-print
  • Hass, C. A., and S. B. Hedges. 1991. Albumin evolution in West Indian frogs of the genus Eleutherodactylus (Leptodactylidae): Caribbean biogeography and a calibration of the albumin immunological clock. J. Zool. (London) 225:413-426. E-print
  • Hedges, S. B. and R. Thomas. 1991. Cryptic species of snakes (Typhlopidae: Typhlops) from the Puerto Rico Bank detected by protein electrophoresis. Herpetologica 47:448-459. E-print
  • Hedges, S. B., and L. R. Maxson. 1991. Pancreatic polypeptide and the sister group of birds. Mol. Biol. Evol. 8:888-891. E-print
  • Hedges, S. B., R. L. Bezy, and L. R. Maxson. 1991. Phylogenetic relationships and biogeography of xantusiid lizards inferred from mitochondrial DNA sequences. Mol. Biol. Evol. 8:767-780. E-print
  • Thomas, R., and S. B. Hedges. 1991. Rediscovery and description of the Hispaniolan lizard Anolis darlingtoni (Sauria, Iguanidae). Carib. J. Sci. 27:90-93. E-print
  • Hedges, S. B. 1991. Electrophoretic and morphological variation in Eleutherodactylus glaphycompus (Anura, Leptodactylidae) of Hispaniola. J. Herpetology 25:10-17. E-print
  • Hedges, S. B., K. D. Moberg, and L. R. Maxson. 1990. Tetrapod phylogeny inferred from 18S and 28S ribosomal RNA sequences and a review of the evidence for amniote relationships. Mol. Biol. Evol. 7:607-633. E-print
  • Hedges, S. B., and K. L. Burnell. 1990. The Jamaican radiation of Anolis (Sauria: Iguanidae): an analysis of relationships and biogeography using sequential electrophoresis. Carib. J. Sci. 26:31-44. E-print
  • Burnell, K. L., and S. B. Hedges. 1990. Relationships of West Indian Anolis (Sauria: Iguanidae): an approach using slow-evolving protein loci. Carib. J. Sci. 26:7-30. E-print
  • Hedges, S. B. 1990. Species concepts and speciation. (Review of Speciation and its Consequences; Otte, D., and J. A. Endler, eds.). Syst. Zool. 39:95-96. E-print
  • Hedges, S. B. 1990. A new species of Phrynopus (Anura, Leptodactylidae) from Peru. Copeia 1990:108-112. E-print
  • Hedges, S. B. 1989. Evolution and biogeography of West Indian frogs of the genus Eleutherodactylus: slow-evolving loci and the major groups. Pp 305-370 in Woods, C. (ed.), Biogeography of the West Indies: Past, Present, and Future. Sandhill Crane Press, Gainesville, Florida. E-print
  • Hedges, S. B. 1989. An island radiation: allozyme evolution in Jamaican frogs of the genus Eleutherodactylus (Anura, Leptodactylidae). Carib. J. Sci. 25:123-147. E-print
  • Thomas, R., and S. B. Hedges. 1989. A new Celestus (Sauria: Anguidae) from the Chaine de la Selle of Haiti. Copeia 1989:882-887. E-print
  • Hedges, S. B., C. A. Hass, and T. K. Maugel. 1989. Physiological color change in snakes. J. Herpetology 23:450-455. E-print
  • Hedges, S. B. 1989. Geographic protein variation in the Jamaican blind snake, Typhlops jamaicensis (Serpentes: Typhlopidae). Caribb. J. Sci. 25:93-96. E-print
  • Hedges, S. B., and R. Thomas. 1989. A new species of Anolis (Sauria: Iguanidae) from the Sierra de Neiba, Hispaniola. Herpetologica 45:330-336. E-print
  • Hedges, S. B., and R. Thomas. 1989. Supplement to West Indian Amphibians and Reptiles: A Checklist. Milwaukee Public Mus. Contr. Biol. Geol. 77:1-11. E-print
  • Hedges. S. B. 1988. A new diminutive frog from Hispaniola (Leptodactylidae, Eleutherodactylus). Copeia 1988(3):636-641. E-print
  • Thomas, R., and S. B. Hedges. 1988. Two new geckos (Sphaerodactylus) from the southern Dominican Republic. Herpetologica 44(1):96-104. E-print
  • Hedges, S. B. 1987. Vocalization and habitat preference of the Jamaican treefrog, Hyla marianae (Anura, Hylidae). Carib. J. Sci. 23:380-384. E-print
  • Hedges, S. B., R. Thomas, and R. Franz. 1987. A new species of Eleutherodactylus (Anura:Leptodactylidae) from the Massif de la Hotte, Haiti. Copeia 1987:943-949. E-print
  • Hedges, S. B., and R. Thomas. 1987. A new burrowing frog from Hispaniola with comments on the inoptatus group of the genus Eleutherodactylus (Anura:Leptodactylidae). Herpetologica 43:269-279. E-print
  • Hedges, S. B. 1986. An electrophoretic analysis of Holarctic hylid frog evolution. Syst. Zool. 35(1):1­21. E-print
  • Hedges, S. B. 1985. The influence of size and phylogeny on life history variation in reptiles: a response to Stearns. Amer. Nat. 126(2):258-260. E-print
  • Busack, S. D., and S. B. Hedges. 1984. Is the peninsular effect a red herring? Amer. Nat. 123(2):266-275. E-print
  • Hedges, S. B. 1982. Caribbean biogeography: implications of recent plate tectonic studies. Syst. Zool. 31(4):518-522. E-print
  • Mitchell, J. C., and S. B. Hedges. 1980. Ambystoma mabeei (Caudata: Ambystomatidae): an addition to the salamander fauna of Virginia. Brimleyana 3:119-121. E-print
  • Zug, G. R., S. B. Hedges, and S. Sunkel. 1979. Variation in reproductive parameters of three neotropical snakes, Coniophanes fissidens, Dipsas catesbyi, and Imantodes cenchoa. Smithsonian Contr. Zool. 300:1-20. E-print
  • Hedges, S. B. 1977. The presence of the scarlet kingsnake, Lampropeltis triangulum elapsoides Holbrook (Reptilia, Serpentes, Colubridae), in the Florida Keys. Herpetol. Rev. 8(4):125-126. E-print

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